This research is supported by the USGS South Florida Priority Ecosystem Studies Program. We are extremely grateful to the South Florida Water Management District (SFWMD) for providing helicopter access to isolated sites. Christopher McVoy (SFWMD) provided key guidance in site selection as well as field assistance. We thank Thomas Sheehan, Bryan Landacre, David Korejwo, and Andrew Lavenburg for assistance with field and laboratory work. William Orem and Lynn Wingard provided constructive comments on earlier versions of the manuscript.